Furthermore, Santos has also called up veteran midfielder Giorgos Karagounis, Borussia Dortmund defender Sokratis Papastathopoulos and Roma's Vasilis Torosidis.
Alexandros Tzorvas (Apollon Smyrnis), Avraam Papadopoulos (Olympiakos) and Kostas Fortounis (Kaiserslautern) are among the players to have been dropped after initially making the provisional squad.
Greece has upcoming friendlies against Portugal, Nigeria and Bolivia before its opening Group C clash with Colombia in Belo Horizonte on June 14. Greece will also meet Cote d'Ivoire and Japan in Brazi.
Squad in full:
Goalkeepers: Panaglotis Glykos (PAOK), Stefanos Kapino (Panathinaikos), Orestis Karnezis (Granada)
Defenders: Jose Holebas, Giannis Maniatis, Kostas Manolas (Olympiakos), Sokratis Papastathopoulos (Borussia Dortmund), Vasilis Torosidis (Roma), Giorgios Tzavellas (PAOK), Loukas Vyntra (Levante), Vangelis Moras (Verona)
Midfielders: Lazaros Christodoulopoulos (Bologna), Ioannis Fetfatzidis (Genoa), Giorgos Karagounis (Fulham), Kostas Katsouranis (PAOK), Panagiotis Kone (Bologna), Andreas Samaris (Olympiakos), Panagiotis Tachtsidis (Torino), Alexandros Tziolis (Kayserispor)
Forwards: Theofanis Gekas (Konyaspor), Kostas Mitroglou (Fulham), Dimitris Salpingidis (PAOK), Georgios Samaras (Celtic)